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The D-retro-inverso design replaces all L-amino acids with D-amino acids and reverses the sequence a structural strategy that preserves the side-chain topology of the original FOXO4 binding region while making the peptide resistant to proteolytic degradation, dramatically extending plasma half-life compared to the natural L-form

However, the observed effects also resulted in significant proliferative action of thyrocytes

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GHKCu limits this by reducing pro-inflammatory cytokines (e.g., IL-6, TNF-) and promoting antioxidant defenses